Dineley, D. & Metcalf, S. GCR Editor: D. Palmer. 1999. Fossil Fishes of Great Britain. Geological Conservation Review Series No. 16. JNCC, Peterborough, ISBN 1 86107 470 0. The original source material for these web pages has been made available by the JNCC under the Open Government Licence 3.0. Full details in the JNCC Open Data Policy
Cruaday Quarry
Highlights
Cruaday Quarry, in the Orkney Islands, provides an exposure of the Sandwick Fish Bed, a famous source of fossil fish specimens. The fish fauna is rich here, including 14 or 15 species, and the site has provided crucial evidence for comparisons with the network mainland fish sites of the same age.
Introduction
Cruaday Quarry
A key figure in the Cruaday story is Professor T.S. Trail of Edinburgh University, who collected many fossil fishes from northern Scotland, particularly Orkney. He corresponded with Agassiz, and in 1832 or 1833 sent him drawings of fossils, and a map showing localities at Skaill, Quoyloo and Breckaness near Stromness (Andrews, 1982). Traill (1834, 1841) described the geology of Orkney and its fossil fishes. The material that he collected, and managed to convey to Edinburgh without loss (see Traill, 1834, p. 646), was sent to Agassiz, who figured type material of new species of fish from Trail's collection from Orkney (Agassiz, 1833–1845), but that material is now lost, presumably because Trail sold his collection to unknown purchasers in 1843 (Andrews, 1982). Clouston (1845) mentioned fossil fishes and plants from several localities in Orkney, and hoped that they would prove the identity of beds with those in Caithness and the Moray Firth. Reference is made to fossil fishes from Ramna Geo on the Admirality Chart for the Orkney Isles for 1850. It was from here that W. Watt obtained most of the fossil fishes which Hugh Miller saw during his visit to the island in 1848 (Wilson et al., 1935). M'Coy (1854) also wrote about material from Orkney, confirming that the Stromness Flagstones were equivalent to the Caithness Flagstones. Many eminent geologists visited the islands and described the geology and palaeontology (Murchison, 1859a, 1859b; Geikie, 1878; Peach and Horne, 1880). Traquair (1894) showed that the fauna of the Achanarras Limestone in Caithness was very similar to that of Stromness, Cromarty and the Moray Firth, and Flett (1898) introduced a stratigraphical scheme for the Orkney Flagstones on the basis of the fish faunas.
Description
Cruaday Quarry is a large excavation on the eastern brow of Cruaday Hill, and exposes 20 m of dark grey calcareous siltstones. The Sandwick Fish Bed is exposed along the western, then north-western, right-angled faces, with a continuous exposure of 450 m (
The Sandwick Fish Bed
Fauna
Acanthodii: Climatiiformes: Diplacanthidae
Diplacanthus crassisimus Duff, 1842 D. sp.
Rhadinacanthus longispinus Agassiz, 1844 = D. perarmatus M'Coy, 1848
Acanthodii: Acanthodiformes: Acanthodidae
Cheiracanthus murchisoni Agassiz, 1835
Mesacanthus sp.
Placodermi: Antiarcha: Pterichyodidae
Pterichthyodes milleri Miller, 1841
Placodermi: Arthrodira: Coccosteidae
Coccosteus cuspidatus Miller, 1841
Placodermi: Arthrodira: Homosteidae
Homosteus milleri Traquair, 1888
Incertae sedis
Palaeospondylus gunni Traquair, 1890
Osteichthyes: Actinopterygii
Cheirolepis trailli Agassiz, 1835
Osteichthyes: Sarcopterygii:
Osteolepidiformes: Osteolepididae
Osteolepis macrolepidotus Agassiz, 1835
?Osteolepis sp. (a large form)
Gyroptychius agassizi Trail, 1841
Glyptolepis leptopterus Agassiz, 1844
Osteichthyes: Sarcopterygii: Dipnoi: Dipteridae
Dipterus valenciennesi Sedgwick and Murchison, 1828
Osteolepis macrolepidotus is common as complete, well-preserved specimens
Trail (1834) was the first to mention the subdivision of the fauna in the Sandwick fish bed. In a quarry near Skail, 'about 100 ft above the level of the sea' he observed that the lower part of the fish bed contained a diverse fauna, whereas the upper part contained mainly fishes 'of a flattened form, with a granular skin, which appear to belong to the family Raja' (i.e. Coccosteus cuspidatus; see also Agassiz, 1835, Tome II, p. 118). Miller (1858) distinguished a three-fold division in the Sandwick Fish Bed, noting 'the lower layer consisting almost exclusively of Dipterians, chiefly osteolepids, the middle layer of acanthodians, of the genera Cheiracanthus and Diplacanthus, and the upper layer of coccosteids, mostly of one species, the 'Coccosteus decipiens'.
Recent studies confirm these ideas in more detail. Trewin (1976) presented a detailed log of the relative abundances of genera through the upper part of the fish bed at Cruaday, relating the pattern and composition of lacustrine varves in museum specimens to a varve by varve section taken from the quarry. He showed that Dipterus is abundant at the base, followed by osteolepids, then acanthodians, and at the top of the fish bed Coccosteus is abundant. Osteolepis and Dipterus do not overlap in distribution with Coccosteus.
Interpretation
Most authors correlate the Sandwick Fish Bed with the Achanarras Fish Bed. The diversity and abundance of the vertebrates must indicate high productivity of the lake waters with annual variations in salinity and organic (algal) productivity; upset on occasion to produce toxic conditions in the lower layers of water with catastrophic consequences for any vertebrate entering. Preservation in such an environment with little physical disturbance was excellent. Large museum collections of fossil fishes have been obtained in the past from other localities in the Cruaday area, Quoyloo, Instabillie, Hooveth and a small quarry on the south side of the Hill of Cruaday. Ramna Geo, a nearby coastal exposure, no longer yields the prolific quantity of material which was found there in the 1840s and 1850s. Less prolific quarries in the Sandwick Fish Bed occurred at Gairsty Farm near Tronston, Glebe Quarry, and at the crossroads near Roadside. 'A small quarry, about 1/4 mile south-west of Bryameadow, on the south of the Burn of Cruaday' was the most productive locality in 1935 (Wilson et al., 1935). All of these localities confirm the areal extent of the conditions described above (see also Jarvik 1950b, 1950c).
Conclusion
Cruaday Quarry is the best site on Orkney showing an Achanarras horizon fish assemblage hence its conservation value. The fauna, consisting of 14 or 15 species of acanthodians, placoderms and osteichthyans, is closely comparable with the fauna from Achanarras itself. Cruaday Hill Quarry has the added advantage, like Achanarras, of recent detailed sedimentological and palaeoenvironmental studies, which enhance understanding of the nature of the great Orcadian Lake. At present,,Cruaday Hill Quarry is the only site in the Sandwick area where a diverse fish assemblage can still be collected. All other sites that were productive in the past are no longer accessible.