Dineley, D. & Metcalf, S. GCR Editor: D. Palmer. 1999. Fossil Fishes of Great Britain. Geological Conservation Review Series No. 16. JNCC, Peterborough, ISBN 1 86107 470 0.

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Headon Hill

[SZ 315 858][SZ 318 862]

(Potential GCR site)

Highlights

Early Cenozoic amphibian fossils are recorded from the epicontinental–continental sequence of this site in the Isle of Wight, Hampshire. They occur in the sands and clays of the Headon Hill Formation (Late Eocene).

Introduction

The Late Eocene (Priabonian) Headon Hill Formation in the type area in the degraded coastal sections of Headon Hill, Isle of Wight, has produced an abundant fauna of turtles, crocodiles, snakes and lizards. Large parts of the section (Figure 15.16) are obscured by mud flows, but the relevant beds may easily be cleared for further excavation.

The Headon Hill Formation between Alum Bay and Totland has been described by Prestwich (1846), White (1921), Stinton (1971), Cray (1973), Daley and Insole (1984) and Insole and Daley (1985). Accounts of the reptilian faunas have been given by Cray (1973), Meszoely and Ford (1976) and Rage and Ford (1980), but there is as yet no complete overview.

Description

A generalized section of the Headon Hill Formation taken from the south-west corner of Headon Hill, based on Cray (1973) and Insole and Daley (1985), is:

Thickness (m)
Cliff End Member (part of 'Upper Headon Beds')
Clays and marls seen to 6.6
Hatherwood Limestone Member (part of 'Upper Headon Beds')
Limestones 2.8
Lignite (Lignite Bed) 0.7
Limestones 2.7
Linstone Chine Member (part of 'Upper Headon Beds')
White and grey sands (Microchoerus Bed at base) 0.8
Colwell Bay Member ('Middle Headon Beds')
Blue-green clays and sands 2.0
Limnaea Limestone 0.2
Blue, green and brown sandy clays (Venus Bed) c. 4.4
Sands, clays and lignites (Neritina Bed) 2.5
Totland Bay Member (Tower Headon Beds')
Limnaea Limestone (How Ledge Limestone) c. 2.0
Marls, clays, sands and lignites 4.6
Limnaea Limestone 0.4
Green clays and pale sands 4.4
Limnaea Limestone 0.8
Blue and green clays 1.0
Limnaea Limestone 0.25
Green sandy clays 0.7
Green clays seen 1.1

In the early 1970s, large collections of reptiles (particularly squamates) and amphibians were obtained by Mr R.L.E. Ford from units in the Totland Bay Member, in particular from Bed HH2 (Bosma, 1974, fig. 9) beneath a unit of hard limestone named the 'How Ledge Limestone', from a series of green-grey clays. Two localities have yielded herpetofaunas from this stratum: in the undercliff at Headon Hill and in Totland Bay. The How Ledge Limestone occurs along the coast between Hatherwood Point and How Ledge, and it appears that the amphibians occur patchily beneath the entire length of the outcrop. The fossils are all represented by disartic-ulated, and frequently abraded and fragmented, elements that indicate considerable predepositional disturbance.

Fauna

The main collection of fossil reptiles from Headon Hill are in the NHM, Museum National d'Histoire Naturelle (MNHN) Paris and the Stuttgarter Museum fur Paldontologie (Ford Collection). The collections include many mammal amd reptile taxa, as well as amphibians.

Anura: Discoglossidae

'discoglossid 2' of Milner et al. (1982)

Anura: Pelobatidae

Eopelobates sp.

Anura: Palaeobatrachidae

Albionbattrachus wightensis Meszoely et al., 1984

Caudata: Salamandridae

salamandrid indet. cf Megalotriton

Interpretation

The interpretation of the Headon Hill section is essentially that given for the Hordle Cliff (q.v.) outcrop — a basin infilling with a final depositional phase above sea level. The land surface was heavily vegetated and supported a considerable vertebrate fauna. For the latter to have been so apparently abundant, there was probably a substantial invertebrate fauna as well, all dependent upon extensive bodies of freshwater. The climate was subtropical or Mediterranean.

Comparison with other localities

Geographically and stratigraphically, the nearest comparable units to the Totland Bay Formation at Headon Hill are the same stratigraphical unit as at Hordle Cliff [SZ 263 923][SZ 273 918] and Wootton Creek, Fishbourne, on the Isle of Wight [SZ 551 927]. In the 'Osborne Beds' the large discoglossid frog 'discoglossid 2' of Milner et al. (1982) is present (Rage and Ford, 1980). All of the amphibians recorded from Headon Hill are known from the directly correlative sequence at Hordle, but there are many genera known from Hordle that are absent on the Isle of Wight (see above), possibly the result of taphonomic differences (Milner et al., 1982). Elements of the Headon Hill fauna are found in a well-preserved assemblage of anurans and caudates from the Oligocene Lower Hamstead Beds exposed along the foreshore at Cranmore Ledge, Bouldnor, on the Isle of Wight ([SZ 370 900][SZ 405 920]; Milner, 1986). This assemblage has yielded 'discoglossids 1 and 2' of Milner et al. (1982), Eopelobates cf. hinschei and the salamanders Salamandra sansaniensis and Chelotriton cf. paradoxus.

The 'Venus' Bed and Oyster Bed of the Middle Headon Beds exposed along the foreshore at Colwell Bay, Isle of Wight [SZ 327 878][SZ 328 881], yield good fish fossils.

Conclusion

The conservation value of the Headon Hill section lies in its great potential for future collecting, since it has been much less exploited than the equivalent-age units at Hordle Cliff (q.v.).

References