Dineley, D. & Metcalf, S. GCR Editor: D. Palmer. 1999. Fossil Fishes of Great Britain. Geological Conservation Review Series No. 16. JNCC, Peterborough, ISBN 1 86107 470 0. The original source material for these web pages has been made available by the JNCC under the Open Government Licence 3.0. Full details in the JNCC Open Data Policy
Melby
Highlights
Two fish beds are present with a fauna close to that at Achanarras and they offer much potential for further study of this northernmost site of this fauna. The Melby Fish Beds at several locations in Shetland have produced excellent specimens of about ten species. These are important since some of them, particularly Pterichthyodes milleri, are index fossils that show that these strata are the same age as those at Achanarras in Caithness and Cruaday in Orkney. This is close to the northernmost point preserved of the great Orcadian Lake (Astin, 1985, 1990).
Introduction
The Melby Fish Beds were discovered by J. Knox during Geological Survey mapping between 1931 and 1934, and they were described by Watson (1934) and Mykura and Phemister (1976). There are two fish beds, separated by 90 m of predominantly fluvial facies rocks. Spore analysis, and the presence of Pterichthyodes milleri (Watson, 1934), confirms that the Melby horizon correlates to the Achanarras horizon (Allen and Marshall, 1981) and is thus of basal Givetian age.
Description
The Melby Fish Beds
At first it was thought that only one fish bed was present (Watson, 1934). The area shows considerable folding, thrusting and faulting, and the fish beds show much lateral variation. Later Mykura and Phemister showed that there were in fact two fish beds
The Lower Melby Fish Bed is exposed at three sites on the coast: Long Rigg
Watson (1934), Mykura and Phemister (1976) and Hemmings (1978) record the following species from the three sites: Coccosteus sp. and Pterichthyodes milleri from Long Rigg, Coccosteus cuspidatus, Dipterus valenciennesi, Glyptolepis cf leptopterus and Gyroptychius agassizi from Ayre of Huxter, and Coccosteus cuspidatus Miller 1841, Dipterus valenciennesi Sedgwick and Murchison, 1828 and Gyroptychius agassizi Trail!, 1841 from Quilva Taing. The faunal differences among these sites respond to lateral lithological variations traceable along the Lower Melby Fish Bed, but also probably reflect collecting bias, because Ayre of Huxter is the only one of the three where collecting is easy.
The Upper Melby Fish Bed varies along strike, containing local lenticular masses of siltstone. It is almost continuously exposed along the top of the cliff from Foglabanks
Two of Watson's (1934) localities are in the Upper Melby Fish Bed (his localities 1 and 3), but it is not clear which sites these are precisely. Locality 1, also known as 'Brough of Huxter' (Hemmings, 1978), is described as 'Shore 1,600 ft W 38° S of Brough of Huxter, and is very close to Pobie Skeo. Fishes occur in carbonate ribs and nodules within grey shales and siltstones', and Watson (1934) recorded Homosteus milleri, ?Coccosteus cuspidatus and Pterichthys sp. ( = P milleri Hemmings 1978), but the original specimens are lost. Locality 3, described as 'shore 2720 ft S 39° W of Brough at Loch of Huxter' is at
Fish species listed by Mykura and Phemister (1976) from the Upper Melby Fish Bed sites include Cheiracanthus sp., ?Coccosteus cuspidatus and Homosteus milleri from Pobie Skeo, Coccosteus cuspidatus and Mesacanthus sp. from the coast north of Matta Taing
Fauna
Placodermi: Arthrodira: Coccosteidae
Coccosteus cuspidatus Miller, 1841
Coccosteus sp.
Placodermi: Arthrodira: Homosteidae
Homosteus milleri Traquair, 1888
Placodermi: Antiarchi: Asterolepidae
Pterichthyodes milleri Miller, 1841
Osteichthyes: Sarcopterygii: Osteolepidae
Dipterus valenciennesi Sedgwick and Murchison, 1828
Acanthodii: Acanthodiformes: Climatiidae
Cheiracanthus sp.
Acanthodii: Acanthodiformes: Acanthodidae
Mesacanthus sp.
Interpretation
The fish beds are thick, pale grey, sandy siltstones and shales with carbonate-rich fish-bearing ribs and nodules and indicate the existence of lacustrine conditions within an otherwise predominantly fluvial environment. The Melby laminites are different from other Orcadian Basin laminites (e.g. at Achanarras; q.v.) in that the elastic laminae often consist of green clay rather than silt, which is probably derived from nearby volcanics (Hall and Donovan, 1978). The nodules themselves indicate a lake margin setting similar in environment to that represented by the Moray Firth nodule beds (Den of Findon, Tynet Burn; q.v.).
The laminites of the Upper Melby Fish Bed at Matta Taing show varved complex sulphide nodules in the lacustrine shales of the fish bed (Hall and Donovan, 1978). Tectonic activity has deformed and fractured these nodules, and they may show cone-in-cone structure and veins. These authors concluded that the complex mineralogical development of the nodules was controlled by lacustrine regression and transgression, and by groundwater enriched because of nearby volcanic activity.
Conclusion
The Melby Fish Beds are the northernmost expression of the Achanarras horizon, and they appear to show a lake-marginal environment, similar to that seen at Den of Findon (q.v.) and Tynet Burn (q.v.). In addition, however, there is strong evidence for local volcanism at Melby. The fish fauna is typical of the Achanarras horizon, as shown particularly by finds of Pterichthyodes milleri, but seems to lack Osteolepis which is so common in Caithness. The sites are worth conserving as the best locations in Shetland showing fish-bearing Orcadian lake sediments, and further collecting is possible from coastal exposures.