Cleal, C.J., Thomas, B.A., Batten, D.J. & Collinson, M.E. 2001. Mesozoic and Tertiary Palaeobotany of Great Britain. Geological Conservation Review Series No. 22, JNCC, Peterborough, ISBN 1 86107 489 1.

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Botton Head

[NZ 596 020]

Introduction

Botton Head is located near Ingelby Greenhow, in the North Yorkshire Moors National Park and is about 4 km from Broughton Bank. This locality in the Aalenian Saltwick Formation is a comparatively recent find. Preliminary investigations have revealed a reasonably large flora with many good gymnosperm reproductive organs. The site has great potential for future studies on gymnosperm evolution and taxonomy.

There is no published account of the palaeobotany of this site, and the following assessment of its significance is based on manuscript notes by Dr Christopher Hill, deposited with fossil plants from the site, in the Natural History Museum, London. Other manuscripts held by the Museum deal with the palynoflora (J.E. John) and megaspores (A. Leitch).

Description

Stratigraphy

The beds, like those at Broughton Bank and Roseberry Topping, belong to the Saltwick Formation. They are composed of claystones, siltstones, sandy siltstones and sandstones overlying the black shales of the Dogger succession (Figure 3.35). The sediments were deposited after the first erosion phase of channel activity in the area and, therefore, yield an interestingly different assemblage from those of the other two sites. There are no marked channel deposits and for some of the time conditions favoured coal formation.

The plant beds are about 2 m thick and extend more than 1 km laterally. The principal bed is above the main sandstone, and is overlain by a micaceous and carbonaceous siltstone with rootlets, followed by a thin layer of dark grey claystone. Like the beds at Broughton Bank and Roseberry Topping, they are near the base of the Hayburn Formation but, unlike the beds at the other two sites, they indicate deposition in an overbank swamp environment with no marked channel association.

Palaeobotany

The complete list of species recovered from this site is given in (Table 3.1). The principal plant bed contains a relatively low abundance of pioneer plants, such as Pachypteris papillosa and ferns, a relatively greater number of bennettites, and some conifers such as Marskea and Bilsdalea. These assemblages are more characteristic of a mature, fairly stable sedimentary environment.

There is considerable lateral variation in the composition of the flora. In some places the grey and grey-brown shales are about 0.15 m thick and incorporate in the top 0.1 m an Equisetum columnare rhizome bed about 200 mm thick.

The coaly layer is rich in bennettites, such as Nilssoniopteris, Pterophyllum, Zamites and Otozamites, together with pieces of Equisetum columnare and Ptilophyllum hirsutum.

Interpretation

A flora such as this, which is rich in bennettites and conifers, is characteristic of a more mature lop set' phase of fluvio-deltaic activity than is found at Broughton Bank and Roseberry Topping. The composition of the coaly layer suggests that it formed by the accumulation of pieces of plants from the surrounding vegetation. The fragmentary pieces of Equisetum columnare must similarly have come from a water-side colony rather than from plants growing on site. The depth of water must have been too great to permit colonization by horsetails.

Conclusions

 The Botton Head flora is dominated by conifers and bennettites. A number of their reproductive organs have yet to be described. The variation in composition offers future possibilities for discovering associations of vegetative and reproductive organs that might indicate natural affinities. A detailed study of the plant fragments and spore content of the coaly layer might also reveal a better understanding of the delta vegetation that surrounded the area of deposition.

References