May, V.J. & Hansom, J.D. 2003. Coastal Geomorphology of Great Britain, Geological Conservation Review Series No. 28. JNCC, Peterborough, ISBN 1 86107 4840. The original source material for these web pages has been made available by the JNCC under the Open Government Licence 3.0. Full details in the JNCC Open Data Policy
Chapter 7 Sandy beaches and dunes — GCR site reports
Introduction
V.J. May
Sandy beaches and their backing dunes are a common feature of the British coast. Although the European Commission's CORINE project recorded 9.6% of the British coast to be sandy beach (European Commission, 1998), this statistic did not include any diff-foot beaches. Sand beaches and dunes occur throughout the British coast, but are concentrated mainly on the northern and western coasts. For example, 75% of coastal dunes, by area, occur north of the Tees and Solway Firth and sand beaches occur in association with dunes and other sandy structures. Sand beaches also commonly form the lower parts of beaches where shingle ridges occur close to high-water mark. They also occur below many cliffs, for example, the chalk around the Isle of Thanet, Kent, the diffs of eastern England from Flamborough Head to Essex, and along much of the Cornish and Welsh coasts, as well as in association with sand cliffs and other strata that yield sand as a major fraction of weathered debris
Since sand supplies from the upper beach are usually required to build sand dunes, the fact that sandy beaches and dunes commonly coexist is unsurprising. However, some sandy beaches are not backed by dunes, mainly owing to limited throughput of sand, an unfavourable wind regime or lack of availability of a site suitable for deposition.
The relationship between sandy cliffs and beaches has typically been described in the context of beach sediment budgets, beach management and coast protection (e.g. Clayton, 1989b; Psuty and Moreira, 1990; Bird, 1996). Extensive sand cliffs (for example at Bournemouth, Dorset, and Culbin, Moray, and, more widely, along the coast of the Algarve in Portugal and much of the coast of California) often have substantial sand beaches at their foot. Much of the supply of sand from sand cliffs in southern and eastern Britain has been reduced in recent years by coastal protection schemes. For example, before they were progressively protected by sea walls during the 20th century, the Bournemouth cliffs (some 11 km in length) produced about 115 000 m3 of sediment annually (of which 80% was coarse enough to stay on the beaches). By the mid 1980s, this supply had fallen to 4000 m3 a−1, coming mostly from the unprotect ed cliffs to the east (Halcrow Maritime, 1999).
A number of writers have argued for a 'systems approach' to sandy beach study, a point also made strongly in respect of cuffed coasts by Brunsden (1973). Such an approach allows each of the influencing factors affecting beach and dune form to be examined in isolation in order to determine its effect. This methodology allows the links between process and form to be better identified. Therefore, much of the investigation of sandy beaches has focused on changes in beach profiles in response to variations in weather conditions, especially wind, and on beach sediment budgets. Long-term trends in beach morphology and the relationship between beach and dune morphology and ecology typify many other studies. However, it is also apparent from the evidence of GCR sites described in this chapter that change in many sand and dune systems is associated with high magnitude/low frequency events superimposed on the more routine processes. Similarly the relationship between many subsystems that make up these features function over different timescales and with different intensities. For example, the sandy beach and dunes at Gibraltar Point, Lincolnshire, comprise many different subsystems, which include nearshore tidal ridges, a ridge-and-runnel foreshore, and a back-shore with arcuate foreshore dune ridges and dune slacks. The spit protects the upper beach ridge sheltering an area of mature (Old Marsh) from New Marsh by a storm beach, which resulted from an occasional extreme event in the evolution of the area. More change occurred in a few hours in 1922 than during years of normal sedimentation; isolation of the storm effects helps in gaining an understanding of the relative importance of both frequent and infrequent events and evolution. This is a theme that is common to many other sand coast GCR sites: Spurn Head on the Holderness cost responded dramatically to a surge in 1849 and both Spurn Head and Gibraltar Point showed different reactions to the 1953 surge. In terms of the development of sub-parallel dune ridges, Gibraltar Point offers considerable contrasts to the GCR site at South Haven Peninsula, Dorset, mostly because of different tidal and wave conditions and differences in sediment supply. In particular, Gibraltar Point lies in a macrotidal and South Haven in a microtidal environment. In both of these sites the processes operating in one subsystem have important repercussions in all of the others.
It is evident from the GCR sites described in this chapter
In the Chalk, sand derives from attrition of flint and from the release of fossil shell fragments from the chalk itself. Elsewhere, sandstone and soft sediment cliffs provide large quantities of sand to their beaches, which may then be transported alongshore. Erosion of the till coast and shallow seabed off Holderness provides very large volumes of sand and gravel annually that are transported both alongshore to form a large sand spit at Spurn Head and into the North Sea. Along the coast of East Anglia, very large volumes of sand and gravel are derived from erosion of till cliffs, but there are also large volumes in offshore banks that result from the offshore transport of longshore sediment. The sand beaches here largely result from the continued throughput of sand. On the more indented coast of the western and northern British Isles, sand beaches are commonly found in embayments where sand cannot escape, and in estuaries and firths, where sand from landward and seaward sources is locally plentiful. Many beaches that are dominated by gravel at the shoreline are also characterized by extensive, sandy, lower beaches. Similarly many beaches formed in heterogeneous materials are sorted locally into sand and gravel for short periods of time and the sand may be blown into sand dunes to the rear of the beach.
Type | Sediment sources | Geomorphological setting | Wind directions | Exemplar GCR sites |
Foreshore dunes | ||||
Spit dunes | Intertidal banks and longshore | On promontories at estuary mouths with near-parallel or radiating ridges and slacks | More common with onshore prevailing and dominant, but not restricted to this | Forvie, Strathbeg, South Haven Penin-sula, Moth Harlech, Holy Island (Goswick and the Snook), Culbin, Morrich More |
Prograding ness dunes | Accretion at ness, possibly with longshore sediment supply from opposite directions alongshore | On open coast | Prevailing and dominant winds from opposite directions (offshore/ onshore) | Winterton Ness, Barry Links, Tentsmuir |
Offshore island dunes | Offshore, longshore and intertidal drying banks | Offshore or barrier islands narrow, subject to washover, often display time- series development in main direction of longshore transport | Can occur with both onshore and offshore prevailing winds | Scolt Head Island, Blakeney Point recurves (North Norfolk Coast), Pembrey (Carmarthen Bay), Culbin, Morrich More |
Hindshore dunes | ||||
Bay dunes | Restricted in longshore direction | Usually at bay head on indented coasts | Prevailing onshore | Dunnet Bay, Luce Sands, Upton and — Gwithian Towans, Tywyn Aberffraw, Oxwich Bay Sandwood, Balta Island, Torrisdale Bay and Invernaver |
Hindshore dune system | Offshore and intertidal | Extensive sandy coasts | Prevailing and dominant winds from the same direction | Braunton Burrows, Newborough Warren, Ainsdale, Holy Island (Ross Links) |
Hindshore sand plains | Offshore, intertidal and beach | Bay-head and low- lying rocky coasts | High wind-speeds that restrict vertical development | Tywyn Aberffraw |
Although this chapter covers sandy beaches as well as dunes, there is little further introduction to beaches that has not been covered in Chapter 5. However, is Marsden Bay, County Durham, is exceptional, where a sandy (and locally mixed sandy gravel) beach lies at the foot of Magnesian limestone cliffs. This was the site of pioneering work on beach mobility in response to variations in wind and waves over 50 years ago (King, 1953), and for that reason is the first site covered in this chapter. For the rest, wide, sandy beaches are usually associated with — and indeed allow the formation of — dunes, but given their varied location, their varying exposure to waves and their range of tidal conditions, they show considerable differences from place to place.
Sand dunes are most likely to be associated with stable and accreting beaches, with a wide upper beach that allows drying and sediment movement by strong winds. A typical example is the west-facing beach of Dunnet Bay in Caithness, a sand trap with onshore winds. Other wide beaches, especially where they are not fully open to the ocean (as around the Irish Sea) and so have waves with more limited fetch, are frequently barred, with ridges and runnels, as at Ainsdale, Lancashire (see GCR site report). Other barred beaches are found at Holkham Bay, North Norfolk, which is a prograding beach, and Braunton Burrows, Devon, as the aerial photograph
Most beaches are more likely to be suffering erosion than progradation (Bird, 1985), and this is certainly true of the UK. The exceptions are in northernmost England (e.g. Holy Island) or parts of Scotland, where postglacial isostatic rebound has offset present-day sea-level rise. As a result, these wide, prograding beaches are backed by some of the largest dune fields in Britain, particularly where sediment was moved onshore during the later part of the Holocene sea-level rise, such as in much of Scotland. It is no surprise that 71% of the dune area of Britain is in Scotland. With the virtual stabilization of sea level, many beaches have lost volume and dune cliffing has become more common throughout Britain. In places, climatic and/or sea-level changes have led to an oscillation between dune cliffing and dune growth on varying timescales, such periodic clang maintaining some dynamic stability via contributions of sand to the fronting beach. In general, present sea-level rise and lack of new sediment means that cuffed dunes are more common than active foredune growth in Britain.
The sandy beaches described in this chapter are only a small sample of the important beach sites included in the coastal geomorphology 'Block' of the GCR, since the great majority of the GCR sites have sandy beaches of one type or another. Chapters 9 and 11 also include descriptions of sandy beach and dune sites where such features are an important part of the coastal geomorphological assemblage. The great depositional sites of Morrich More in the Dornoch Firth and Culbin in the Moray Firth, the Northumbrian coast around Holy Island, the North Norfolk coast and Rhossili Bay (Carmarthen Bay GCR site) all provide unmodified, dynamic examples of some of the finest sandy beaches to be found in the UK.
Coastal dunes
There are over 295 separate coastal dune sites around Great Britain (shown on the small-scale map in
Site | Main features | Other features | Present-day sediment sources | |
Marsden Bay | Beach phases | Cliffs and stacks | Local cliff erosion — small | 4.2 |
South Haven Peninsula | Shore-parallel dune ridges, originating from the 16th century, slacks, sand-spit | Relict and active cliffs., caves, rock platform | Longshore — restricted Offshore — significant | 1.5 |
Upton and Gwithian Towans | Climbing dunes, exhumed bedrock base | Stacks | Offshore — restricted | 5.8 |
Braunton Burrows | Large dune field, parabolic dunes, slacks | Ridge and runnel | Intertidal and estuarine | 7.3 |
Oxwich Bay | Bay-head beach and dunes | Cliffs and emerged platform | Offshore — limited | 8.2 |
Tywyn Aberffraw | Sand plain, isolated parabolic dunes shore- parallel linear dunes | Offshore, probably in deficit | 4.7 | |
Ainsdlale | Large dune field, slacks, ridge and runnel, long dated history | Offshore — limited — in deficit | 8.3 | |
Luce Sands | Bay-head dunes | Holocene emerged gravel ridges | Onshore and longshore — significant | 5.6 |
Sandwood Bay | Dynamic beach-dune complex, climbing dunes | Gravel-cored bar, blowouts | Offshore and recycled — limited | 4.2 |
Dunnet Bay | Bay-head dunes and sand plain | Blowouts | Offshore — limited | 4.0 |
Baba Island | Climbing dunes | Beach-dune-grassland continuum | Local — limited | 1.9 |
Strathbeg | Shore-parallel dune ridges, large blowouts | Holocene emerged gravel ridges | Longshore — restricted, loch outlet source | 3.3 |
Forvie | Shore-parallel dune ridges, originally moved as waves northwards | Longshore — cycled from estuary | 3.1 | |
Barry Links | Foreland sand plain, linear parabolic dunes | Estuarine, longshore — limited | 4.4 | |
Tentsmuir | Shore-parallel dune ridges-intertidal sands | Estuarine and longshore — significant | 4.4 | |
Torrisdale and Invernaver | Beach-dune, hill-top dunes, glaciofluvial terraces | Archaeological context | Offshore and fluvial recycled — now limited | 4.0 |
Morrich More | Shore-parallel beaches and dunes: sandplain | Holocene beaches and cliffs | Offshore — restricted | 4.3 |
Culbin | Shore-parallel dunes, large dune field now stabilized by forest | Holocene emerged gravel ridges and spits | Longshore -restricted, offshore — limited | 3.6 |
East Head | Small spit-based dunes | Intertidal | 3.4 | |
Holy Island | Dune field, spits, barrier beach | Cliffs, Holocene saltmarsh, intertidal mudflats | Longshore, offshore — significant | 4.1 |
Dawlish Warren | Parallel spit-based linear dunes | Recurved spit | Intertidal and possibly estuarine In deficit | 4.1 |
North Norfolk Coast | Major mainly linear dunes | Spits, barrier beach | Longshore and offshore | 6A-4.7 |
Morfa Harlech | Linear shore-parallel dunes | Longshore — restricted, estuarine | 4.5 | |
Morfa Dyffryn | Linear shore-parallel dunes, blowouts, dunes invading slacks | Longshore — restricted, offshore | 4.3 | |
Winterton Ness | Linear dunes on cuspate foreland | Longshore | 2.6 | |
Ynyslas | Spit-based dunes | Longshore — restricted, estuarine | 4.3 | |
Carmarthen Bay | ||||
Pendine | Shore-parallel linear dunes | Offshore, estuarine to distal end | 8.0 | |
Pembrey | Large dune field, spit-based linear dunes | Offshore and estuarine | 8.0 | |
Whitford spit | Estuary-mouth spit | Longshore, drying intertidal | 8.0 | |
Laugharne Burrows | Cliff-top dunes | Local redistribution, drying intertidal | 8.0 | |
Newborough Warren and Morfa Dinlle | Major dune field, parabolic and linear dunes, spit, tied island and slacks | Saltmarsh | Offshore and estuarine | 4.7 |
Dune location | CaCO3
(%) |
Median
grain size (Phi) |
Culbin | 0.0 | 2.0 |
South Haven Peninsula | 0.015 | ? |
Lossiemouth | 0.26 | 2.0 |
Tentsmuir | 0.4 | 2.5 |
Luce Sands | 0.5 | 2.4 |
Forvie | 0.55 | 1.9 |
Buddon Ness (Barry Links) | 1.0 | 2.0 |
Walney Island | 1.51 | 2.21 |
Morfa Dyffryn | 3.34 | 2.31 |
Ainsdale | 3.57 | 2.13 |
Invernaver | 3.8 | 1.9 |
Morfa Harlech | 3.96 | 2.13 |
Newborough Warren | 4.56 | 2.50 |
Ynyslas | 4.98 | 2.29 |
Strathbeg | 7.86 | 2.0 |
Rattray (Strathbeg site) | 9.10 | 1.9 |
Laugharne (Pendine)* | 11.15 | 2.40 |
Morrich More | 12.0 | 2.4 |
Pembrey* | 12.04 | 2.33 |
Oxwich Bay | 12.45 | 1.93 |
Tywyn Aberffraw | 13.20 | 2.47 |
Llangennith* | 15.65 | 1.63 |
Braunton Burrows | 19.59 | 2.13 |
Dunnet Bay | 20.4 | 1.7 |
Dunbar | 20.4 | 1.5 |
Westward Ho! | 21.79 | 2.45 |
Machir, Islay | 33.6 | 2.2 |
Mangersta, Lewis | 38 | 1.4 |
Luskentyre, Harris | 44 | 2.0 |
Tràigh na Berie, Lewis | 47 | 2.4 |
St. Ninian's Tombolo, Shetland | 47.5 | 2.0 |
Balnakiel | 52.0 | 1.8 |
Hayle (Upton and Gwithian Towans) | 56.80 | 1.56 |
Loch Gruinart, Islay | 59.0 | 2.1 |
Eoligarry, Barra | 80.0 | 2.0 |
Ardivachar, South Uist | 84.0 | 1.7 |
Balta Island, Shetland | 95.5 | 1.8 |
*Camarthen Bay |
Most British dune systems originated when substantial seabed deposits were moved onshore during the early and middle part of the Holocene Epoch and began to be deposited dose to their present locations from about 6500 years BP In some areas where the sea-level history is more complex, such as in the Western Isles of Scotland, the arrival of dune sands first began about 8700 years BP and may have been non-synchronous between sites (see Chapter 9; Hansom and Angus, 2001). Dune systems such as those at Ainsdale and Braunton Burrows can be shown to have developed over the past six millennia, especially from the evidence of preserved peat associated with dune slacks and larger wetlands that developed shorewards of the coastal beaches. In contrast, other dunes are more recent, for example at South Haven Peninsula the dunes have formed since the 16th century. Some dunes, for example at Culbin, Moray, Newborough Warren on the Isle of Angelsey, and Hayle and Upton and Gwithian Towans, Cornwall, have migrated inland covering buildings and farmland. British dunes tend to be located:
- in areas of high tidal range,
- where prevailing winds provide the main means of landward aeolian transport, and
- in association with estuary mouths dominated by large sandy sediment loads or at the heads of inlets and bays,
- on north-eastern coasts, where strong winds from the north and east provide the means for landward aeolian transport e.g. the coasts between Aberdeen and Fraserburgh and Northumberland.
Narrow, linear-dune systems occur along eastern coasts that are associated with sandy estuaries or high tidal ranges, but the size of the dunes is generally much less than those of the exposed and windy western coasts, even though the intertidal sandy area may be very extensive.
There are few significant dunes on the eastern coast of England, apart from the dunes around Holy Island, Northumberland, and along the Lincolnshire and north Norfolk coasts. Between the Tees and the Tamar there are 24 dune sites (c. 8%) and between the Tamar and the Mull of Galloway 67 dune sites (c. 23%). The remaining 204 (c. 69%) sites lie along the coast of Scotland and the English coast north of the Tees. The largest area of dunes is in north-west Scotland, particularly in the Outer Hebrides where machair predominates (Ritchie and Mather, 1984; Dargie, 2000; see Chapter 9). Of 43 nationally important sand dune sites, only six lie on the south or east coast (Doody, 1985).
Location | Calcium carbonate | content of dunes | pH | |
Foredunes | Main dunes | Foredunes | Main dunes | |
South Haven Peninsula | 0.015 | 0.01 | 7.0 | 3.6 |
Southport (near Ainsdale) | 6.0 | 0.2 | 8.2 | 5.5 |
Braunton Burrows | 20.0 | 8.5 | 9.05 | 8.2 |
Blakeney Point, North Norfolk Coast | 0.6 | 0.02 | 7.3 | 4.2 |
The foredunes around the coast of England and Wales are notable for their generally low calcium carbonate content
On much of the southern coast of Britain, sand was in plentiful supply for dune building at the end of the main Holocene rise when sea level attained present levels about 6500 years BP In recent centuries, however, the supply of sand has diminished significantly and erosional conditions generally prevail.
In England, few southern or eastern dunes are accreting, the most important exceptions being at Holy Island and South Haven Peninsula, and even the latter is affected by erosion of its older southern beach and dunes. In contrast, on western and northern coasts, dunes are common features, reflecting the combination of plentiful sand supplies mainly from the seabed in the past, but also from upland river catchments, and the effects of prevailing onshore winds. However, many are now affected by wave erosion of their fronts either by occasional storms or by long-term changes in sea-level and storminess, together with reduced sediment supply. Prior to 6500 years BP sand supply for dune building was plentiful, but it is now much reduced, and, as a result, frontal dune erosion is commonplace (Hansom, 1988; Hansom and Angus, 2001).
The conservation value of sandy beaches and dunes
Dunes are geomorphologically important because of:
- their natural dynamism and the relationship with their ecology
- their role in preserving and then exhuming Holocene sedimentary sequences and
- their role in coast protection.
The selected GCR sites
Most dune systems around the British coast are complex, and very few have individual isolated stable dunes within them. English east coast dunes are generally narrow, have only limited periods of onshore winds, and lack large and constant sand supplies. Many of those on the west coast lie upon bedrock surfaces of varying height and so lack the level foundations of sand plains. They also have usually had ample supplies of sand in the past that have produced a complex dune topography in which 'dunes are at many stages of development and sand is transferred from erosional phases to depositional ones (for example at Newborough Warren and Morfa Dyffryn). Tywyn Aberffraw is an important member of the network of dune systems because of its relatively limited sediment supply and restricted development of dunes. In this respect it contrasts especially strongly with its neighbour at Newborough Warren.
In this chapter the site reports are ordered in a clockwise fashion starting with the Marsden Bay GCR site.
Dunes and sandy beaches as biological SSSIs and Special Areas of Conservation (SACs)
In Chapter 1, it was emphasized that the SSSI site series is constructed both from areas nationally important for wildlife, and GCR sites. An SSSI may be established solely for its geology/ geomorphology, or its wildlife/habitat, or it may comprisc a 'mosaic' of biological and GCR sites that may be adjacent, partly overlap, or be coincident. There are a number of sand dune and beach sites that are crucially important to the natural heritage of Britain that are notified as SSSIs primarily for their wildlife value, but implicitly will contain interesting coastal geomorphology features that are not included independently in the GCR because of the 'minimum number' criterion of the GCR rationale (see Chapter 1). These sites are not described in the present geomorphologically focused volume.
SAC name | Local authority | Dune habitat extent (ha) |
Barry Links | Angus | 447.6 |
Braunton Burrows | Devon | 767.5 |
Carmarthen Bay Dunes/Twyni Bae Caerfyrddin | Abertawe/ Swansea; Caerfyrddin/ Carmarthenshire | 780.2 |
Coll Machair | Argyll and Bute | 409.0 |
Culbin Bar | Highland; Moray | 612.9 |
Dawlish Warren | Devon | 28.2 |
Dee Estuary/ Aber Dyfrdwy* | Cheshire; Fflint/ Flintshire; Wirral | 4.0 |
Dornoch Firth and Morrich More | Highland | 974.4 |
Dorset Heaths (Purbeck and Wareham) and Studland Dunes | Dorset | 95.9 |
Drigg Coast | Cumbria | 519.8 |
Durness | Highland | 386.7 |
Humber Estuary* | City of Kingston upon Hull; East Riding of Yorkshire; Lincolnshire; North East Lincolnshire; North Lincolnshire | 529.0 |
Invernaver | Highland | 54.2 |
Kenfig/ Cynffig | Pen-y-bont ar Ogwr/ Bridgend | 673.8 |
Limestone Coast of South West Wales/ Arfordir Calchfaen de Orllewin Cymru | Abertawe/ Swansea; Penfro/ Pembrokeshire | 397.1 |
Monach Islands | Western Isles / Na h-Eileanan an lar | 215.1 |
Morecambe Bay | Cumbria; Lancashire | 220.5 |
Morfa Harlech a Morfa Dyffryn | Gwynedd | 228.6 |
North Norfolk Coast | Norfolk | 387.3 |
North Northumberland Dunes | Northumberland | 1078.6 |
North Uist Machair | Western Isles / Na h-Eileanan an lar | 963.3 |
Oldshoremore and Sandwood | Highland | 165.3 |
Penhale Dunes | Cornwall | 422.4 |
Saltfleetby-Theddlethorpe Dunes and Gibraltar Point | Lincolnshire | 265.6 |
Sands of Forvie | Aberdeenshire | 469.7 |
Sandwich Bay | Kent | 258.3 |
Sefton Coast | Sefton | 1072.7 |
Solent Maritime | City of Portsmouth; City of Southampton; Hampshire; Isle of Wight; West Sussex | 113.2 |
Solway Firth | Cumbria; Dumfries and Galloway | 32.6 |
South Uist Machair | Western Isles / Na h-Eileanan an lar | 545.7 |
Tiree Machair | Argyll and Bute | 237.4 |
Torrs Warren-Luce Sands | Dumfries and Galloway | 819.5 |
Winterton-Horsey Dunes | Norfolk | 44.7 |
Y Twyni o Abermenai i Aberffraw/ Abermenai to Aberffraw Dunes | Gwynedd; Ynys Mon/ Isle of Anglesey | 672.3 |
* Possible SAC not yet submitted to EC. | Bold type indicates a coastal GCR interest within the site. |
The importance of dunes as areas of national ecological significance was recognized and described by Tansley (1939, 1945) and Steers (1946a, 1953a). Soon after the Nature Conservancy was established in 1949, it designated a number of major dunes as National Nature Reserves, including Braunton Burrows, Newborough Warren, Ainsdale and Holy Island. The Nature Conservation Review (Ratcliffe, 1977) confirmed the great importance of dunes as part of the network of nationally significant sites.
In addition to being protected through the SSSI system for their national importance, certain types of dune are Habitats Directive Annex I habitats, eligible for selection as Special Areas of Conservation (see Chapter 1). The Directive identifies a suite of dune vegetation types (see below), representing the succession from dune initiation to mature, stable dune habitat. Collectively, these types encompass almost the full range of coastal dune habitats present in the UK.
Dune SAC site selection rationale
The sites are, for the most part, the most extensive examples in the UK and have the best conserved structure and function, demonstrating transitions between Annex I types, while also representing the range of geographic and ecological variation of each habitat type.
- Embryonic shifting dune vegetation exists in a highly dynamic state and is dependent on the continued operation of physical processes at the dune/beach interface. It is the first type of vegetation to colonize areas of incipient dune formation at the top of a beach.
- Shifting dunes along the shoreline with Ammophila arenaria ('white dunes') encompass most of the vegetation of unstable dunes where there is active sand movement. Under these conditions sand-binding marram A. arenaria is always a prominent feature of the vegetation and is usually dominant.
- Fixed dune vegetation occurs mainly on the largest dune systems, being those that have the width to allow it to develop. It typically occurs inland of the zone dominated by mar-ram Ammophila arenaria on coastal dunes, and represents the vegetation that replaces marram as the dune stabilizes and the organic content of the sand increases.
- Decalcified fixed dunes with crowberry Empetrum nigrumrepresent the later, more mature, stages of the successional sequence characteristic of sand dunes. Exposure to rainfall over long periods means that there is leaching of the surface layers, causing a loss of calcium carbonate and increased soil acidity.
- Atlantic decalcified fixed dunes (Calluno-Ulicetea)occur on mature, stable dunes where the initial calcium carbonate content of the dune sand is low. The surface soil layers rapidly lose their remaining calcium carbonate through leaching, and become acidified.
- Dunes with Hippophae rbamnoidescomprise scrub vegetation on more-or-less stable sand dunes in which sea-buckthorn is abundant.
- Dunes with Salix repens ssp. argentea, where creeping willow is dominant, forming prominent, low scrubby growth.
- Humid dune slacks are low-lying areas within dune systems that are seasonally flooded and where nutrient levels are low. Dune slacks are often rich in plant species.
- Coastal dunes with juniperJuniperus spp. comprises common juniper scrub in a variety of dune situations.
- Machair — see Chapter 9 of the present volume.